"Nature," November 11th, 1880, page 32.)

Down, November 5th, 1880.

Sir Wyville Thomson and Natural Selection.

I am sorry to find that Sir Wyville Thomson does not understand the principle of Natural Selection, as explained by Mr. Wallace and myself. If he had done so, he could not have written the following sentence in the Introduction to the Voyage of the "Challenger": "The character of the abyssal fauna refuses to give the least support to the theory which refers the evolution of species to extreme variation guided only by Natural Selection." This is a standard of criticism not uncommonly reached by theologians and metaphysicians, when they write on scientific subjects, but is something new as coming from a naturalist. Professor Huxley demurs to it in the last number of "Nature"; but he does not touch on the expression of extreme variation, nor on that of evolution being guided only by Natural Selection. Can Sir Wyville Thomson name any one who has said that the evolution of species depends only on Natural Selection? As far as concerns myself, I believe that no one has brought forward so many observations on the effects of the use and disuse of parts, as I have done in my "Variation of Animals and Plants under Domestication"; and these observations were made for this special object. I have likewise there adduced a considerable body of facts, showing the direct action of external conditions on organisms; though no doubt since my books were published much has been learnt on this head. If Sir Wyville Thomson were to visit the yard of a breeder, and saw all his cattle or sheep almost absolutely true--that is, closely similar, he would exclaim: "Sir, I see here no extreme variation; nor can I find any support to the belief that you have followed the principle of selection in the breeding of your animals." From what I formerly saw of breeders, I have no doubt that the man thus rebuked would have smiled and said not a word. If he had afterwards told the story to other breeders, I greatly fear that they would have used emphatic but irreverent language about naturalists.

(301/2. The following is the passage omitted by the advice of Huxley: see his "Life and Letters," II., page 14:--

"Perhaps it would have been wiser on my part to have remained quite silent, like the breeder; for, as Prof. Sedgwick remarked many years ago, in reference to the poor old Dean of York, who was never weary of inveighing against geologists, a man who talks about what he does not in the least understand, is invulnerable.")

LETTER 302. TO G.J. ROMANES.

(302/1. Part of this letter has been published in Mr. C. Barber's note on "Graft-Hybrids of the Sugar-Cane," in "The Sugar-Cane," November 1896.)

Down, January 1st, 1881.

I send the MS., but as far as I can judge by just skimming it, it will be of no use to you. It seems to bear on transitional forms. I feel sure that I have other and better cases, but I cannot remember where to look.

I should have written to you in a few days on the following case. The Baron de Villa Franca wrote to me from Brazil about two years ago, describing new varieties of sugar-cane which he had raised by planting two old varieties in apposition. I believe (but my memory is very faulty) that I wrote that I could not believe in such a result, and attributed the new varieties to the soil, etc. I believe that I did not understand what he meant by apposition. Yesterday a packet of MS. arrived from the Brazilian Legation, with a letter in French from Dr. Glass, Director of the Botanic Gardens, describing fully how he first attempted grafting varieties of sugar-cane in various ways, and always failed, and then split stems of two varieties, bound them together and planted them, and then raised some new and very valuable varieties, which, like crossed plants, seem to grow with extra vigour, are constant, and apparently partake of the character of the two varieties. The Baron also sends me an attested copy from a number of Brazilian cultivators of the success of the plan of raising new varieties. I am not sure whether the Brazilian Legation wishes me to return the document, but if I do not hear in three or four days that they must be returned, they shall be sent to you, for they seem to me well deserving your consideration.

Charles Darwin

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