In this discussion, we are considering only formations containing the remains of marine animals, which, as before mentioned, live, with some exceptions within (most of them much within) depths of 100 fathoms. How, then, can a thick and widely extended formation be accumulated, which shall include such organic remains? First, let us take the case of the bed of the sea long remaining at a stationary level: under these circumstances it is evident that CONCHIFEROUS strata can accumulate only to the same thickness with the depth at which the shells can live; on gently inclined coasts alone can they accumulate to any considerable width; and from the want of superincumbent pressure, it is probable that the sedimentary matter will seldom be much consolidated: such formations have no very good chance, when in the course of time they are upraised, of long resisting the powers of denudation. The chance will be less if the submarine surface, instead of having remained stationary, shall have gone on slowly rising during the deposition of the strata, for in this case their total thickness must be less, and each part, before being consolidated or thickly covered up by superincumbent matter, will have had successively to pass through the ordeal of the beach; and on most coasts, the waves on the beach tend to wear down and disperse every object exposed to their action. Now, both on the south-eastern and western shores of South America, we have had clear proofs that the land has been slowly rising, and in the long lines of lofty cliffs, we have seen that the tendency of the sea is almost everywhere to eat into the land. Considering these facts, it ceases, I think, to be surprising, that extensive recent conchiferous deposits are entirely absent on the southern and western shores of America.

Let us take the one remaining case, of the bed of the sea slowly subsiding during a length of time, whilst sediment has gone on being deposited. It is evident that strata might thus accumulate to any thickness, each stratum being deposited in shallow water, and consequently abounding with those shells which cannot live at great depths: the pressure, also, I may observe, of each fresh bed would aid in consolidating all the lower ones. Even on a rather steep coast, though such must ever be unfavourable to widely extended deposits, the formations would always tend to increase in breadth from the water encroaching on the land. Hence we may admit that periods of slow subsidence will commonly be most favourable to the accumulation of CONCHIFEROUS deposits, of sufficient thickness, extension, and hardness, to resist the average powers of denudation.

We have seen that at an ancient tertiary epoch, fossiliferous deposits were extensively deposited on the coasts of South America; and it is a very interesting fact, that there is evidence that these ancient tertiary beds were deposited during a period of subsidence. Thus, at Navidad, the strata are about eight hundred feet in thickness, and the fossil shells are abundant both at the level of the sea and some way up the cliffs; having sent a list of these fossils to Professor E. Forbes, he thinks they must have lived in water between one and ten fathoms in depth: hence the bottom of the sea on which these shells once lived must have subsided at least 700 feet to allow of the superincumbent matter being deposited. I must here remark, that, as all these and the following fossil shells are extinct species, Professor Forbes necessarily judges of the depths at which they lived only from their generic character, and from the analogical distribution of shells in the northern hemisphere; but there is no just cause from this to doubt the general results. At Huafo the strata are about the same thickness, namely, 800 feet, and Professor Forbes thinks the fossils found there cannot have lived at a greater depth than fifty fathoms, or 300 feet. These two points, namely, Navidad and Huafo, are 570 miles apart, but nearly halfway between them lies Mocha, an island 1,200 feet in height, apparently formed of tertiary strata up to its level summit, and with many shells, including the same Turritella with that found at Huafo, embedded close to the level of the sea.

Charles Darwin

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