This summer will make the 20th year (!) since I opened my first note-book, on the question how and in what way do species and varieties differ from each other. I am now preparing my work for publication, but I find the subject so very large, that though I have written many chapters, I do not suppose I shall go to press for two years. I have never heard how long you intend staying in the Malay Archipelago; I wish I might profit by the publication of your Travels there before my work appears, for no doubt you will reap a large harvest of facts. I have acted already in accordance with your advice of keeping domestic varieties, and those appearing in a state of nature, distinct; but I have sometimes doubted of the wisdom of this, and therefore I am glad to be backed by your opinion. I must confess, however, I rather doubt the truth of the now very prevalent doctrine of all our domestic animals having descended from several wild stocks; though I do not doubt that it is so in some cases. I think there is rather better evidence on the sterility of hybrid animals than you seem to admit: and in regard to plants the collection of carefully recorded facts by Kolreuter and Gaertner (and Herbert,) is ENORMOUS. I most entirely agree with you on the little effects of "climatal conditions," which one sees referred to ad nauseam in all books: I suppose some very little effect must be attributed to such influences, but I fully believe that they are very slight. It is really IMPOSSIBLE to explain my views (in the compass of a letter), on the causes and means of variation in a state of nature; but I have slowly adopted a distinct and tangible idea,--whether true or false others must judge; for the firmest conviction of the truth of a doctrine by its author, seems, alas, not to be the slightest guarantee of truth!...

CHARLES DARWIN TO J.D. HOOKER. Moor Park, Saturday [May 2nd, 1857].

My dear Hooker,

You have shaved the hair off the Alpine plants pretty effectually. The case of the Anthyllis will make a "tie" with the believed case of Pyrenees plants becoming glabrous at low levels. If I DO find that I have marked such facts, I will lay the evidence before you. I wonder how the belief could have originated! Was it through final causes to keep the plants warm? Falconer in talk coupled the two facts of woolly Alpine plants and mammals. How candidly and meekly you took my Jeremiad on your severity to second-class men. After I had sent it off, an ugly little voice asked me, once or twice, how much of my noble defence of the poor in spirit and in fact, was owing to your having not seldom smashed favourite notions of my own. I silenced the ugly little voice with contempt, but it would whisper again and again. I sometimes despise myself as a poor compiler as heartily as you could do, though I do NOT despise my whole work, as I think there is enough known to lay a foundation for the discussion on the origin of species. I have been led to despise and laugh at myself as a compiler, for having put down that "Alpine plants have large flowers," and now perhaps I may write over these very words, "Alpine plants have small or apetalous flowers!"...

CHARLES DARWIN TO J.D. HOOKER. Down, [May] 16th [1857].

My dear Hooker,

You said--I hope honestly--that you did not dislike my asking questions on general points, you of course answering or not as time or inclination might serve. I find in the animal kingdom that the proposition that any part or organ developed normally (i.e., not a monstrosity) in a species in any HIGH or UNUSUAL degree, compared with the same part or organ in allied species, tends to be HIGHLY VARIABLE. I cannot doubt this from my mass of collected facts. To give an instance, the Cross-bill is very abnormal in the structure of its bill compared with other allied Fringillidae, and the beak is EMINENTLY VARIABLE. The Himantopus, remarkable from the wonderful length of its legs, is VERY variable in the length of its legs. I could give MANY most striking and curious illustrations in all classes; so many that I think it cannot be chance.

Charles Darwin

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