n from the forefather to his descendants of dormant gemmules, which occasionally become developed under certain known or unknown conditions. Each animal and plant may be compared with a bed of soil full of seeds, some of which soon germinate, some lie dormant for a period, whilst others perish. When we hear it said that a man carries in his constitution the seeds of an inherited disease, there is much truth in the expression. No other attempt, as far as I am aware, has been made, imperfect as this confessedly is, to connect under one point of view these several grand classes of facts. An organic being is a microcosm--a little universe, formed of a host of self-propagating organisms, inconceivably minute and numerous as the stars in heaven.

CHAPTER 2.XXVIII.

CONCLUDING REMARKS.

DOMESTICATION. NATURE AND CAUSES OF VARIABILITY. SELECTION. DIVERGENCE AND DISTINCTNESS OF CHARACTER. EXTINCTION OF RACES. CIRCUMSTANCES FAVOURABLE TO SELECTION BY MAN. ANTIQUITY OF CERTAIN RACES. THE QUESTION WHETHER EACH PARTICULAR VARIATION HAS BEEN SPECIALLY PREORDAINED.

As summaries have been added to nearly all the chapters, and as, in the chapter on pangenesis, various subjects, such as the forms of reproduction, inheritance, reversion, the causes and laws of variability, etc., have been recently discussed, I will here only make a few general remarks on the more important conclusions which may be deduced from the multifarious details given throughout this work.

Savages in all parts of the world easily succeed in taming wild animals; and those inhabiting any country or island, when first visited by man, would probably have been still more easily tamed. Complete subjugation generally depends on an animal being social in its habits, and on receiving man as the chief of the herd or family. In order that an animal should be domesticated it must be fertile under changed conditions of life, and this is far from being always the case. An animal would not have been worth the labour of domestication, at least during early times, unless of service to man. From these circumstances the number of domesticated animals has never been large. With respect to plants, I have shown in the ninth chapter how their varied uses were probably first discovered, and the early steps in their cultivation. Man could not have known, when he first domesticated an animal or plant, whether it would flourish and multiply when transported to other countries, therefore he could not have been thus influenced in his choice. We see that the close adaptation of the reindeer and camel to extremely cold and hot countries has not prevented their domestication. Still less could man have foreseen whether his animals and plants would vary in succeeding generations and thus give birth to new races; and the small capacity of variability in the goose has not prevented its domestication from a remote epoch.

With extremely few exceptions, all animals and plants which have been long domesticated have varied greatly. It matters not under what climate, or for what purpose they are kept, whether as food for man or beast, for draught or hunting, for clothing or mere pleasure,--under all these circumstances races have been produced which differ more from one another than do the forms which in a state of nature are ranked as different species. Why certain animals and plants have varied more under domestication than others we do not know, any more than why some are rendered more sterile than others under changed conditions of life. But we have to judge of the amount of variation which our domestic productions have undergone, chiefly by the number and amount of difference between the races which have been formed, and we can often clearly see why many and distinct races have not been formed, namely, because slight successive variations have not been steadily accumulated; and such variations will never be accumulated if an animal or plant be not closely observed, much valued, and kept in large numbers.

The fluctuating, and, as far as we can judge, never-ending variability of our domesticated productions,--the plasticity of almost their whole organisation,- -is one of the most important lessons which we learn from the numerous details given in the earlier chapters of this work.

Charles Darwin

All Pages of This Book