'Annal. des Sc. Nat.' 3rd series, Bot. tome 6 page 189.) alludes to the extreme vigour of some of the crossed fruit- trees raised by Sageret.

By crossing reciprocally the tallest and shortest peas, Knight (17/53. 'Philosophical Transactions' 1799 page 200.) says: "I had in this experiment a striking instance of the stimulative effects of crossing the breeds; for the smallest variety, whose height rarely exceeded two feet, was increased to six feet: whilst the height of the large and luxuriant kind was very little diminished." Mr. Laxton gave me seed-peas produced from crosses between four distinct kinds; and the plants thus raised were extraordinarily vigorous, being in each case from one to two or three feet taller than the parent-forms growing close alongside them.

Wiegmann (17/54. 'Ueber die Bastarderzeugung' 1828 s. 32, 33. For Mr. Chaundy's case see Loudon's 'Gardener's Mag.' volume 7 1831 page 696.) made many crosses between several varieties of cabbage; and he speaks with astonishment of the vigour and height of the mongrels, which excited the amazement of all the gardeners who beheld them. Mr. Chaundy raised a great number of mongrels by planting together six distinct varieties of cabbage. These mongrels displayed an infinite diversity of character; "But the most remarkable circumstance was, that, while all the other cabbages and borecoles in the nursery were destroyed by a severe winter, these hybrids were little injured, and supplied the kitchen when there was no other cabbage to be had."

Mr. Maund exhibited before the Royal Agricultural Society (17/55. 'Gardener's Chronicle' 1846 page 601.) specimens of crossed wheat, together with their parent varieties; and the editor states that they were intermediate in character, "united with that greater vigour of growth, which it appears, in the vegetable as in the animal world, is the result of a first cross." Knight also crossed several varieties of wheat (17/56. 'Philosoph. Transact.' 1799 page 201.), and he says "that in the years 1795 and 1796, when almost the whole crop of corn in the island was blighted, the varieties thus obtained, and these only, escaped in this neighbourhood, though sown in several different soils and situations."

Here is a remarkable case: M. Clotzsch (17/57. Quoted in 'Bull. Bot. Soc. France' volume 2 1855 page 327.) crossed Pinus sylvestris and nigricans, Quercus robur and pedunculata, Alnus glutinosa and incana, Ulmus campestris and effusa; and the cross-fertilised seeds, as well as seeds of the pure parent-trees, were all sown at the same time and in the same place. The result was, that after an interval of eight years, the hybrids were one-third taller than the pure trees!

The facts above given refer to undoubted varieties, excepting the trees crossed by Clotzsch, which are ranked by various botanists as strongly-marked races, sub-species, or species. That true hybrids raised from entirely distinct species, though they lose in fertility, often gain in size and constitutional vigour, is certain. It would be superfluous to quote any facts; for all experimenters, Kolreuter, Gartner, Herbert, Sageret, Lecoq, and Naudin, have been struck with the wonderful vigour, height, size, tenacity of life, precocity, and hardiness of their hybrid productions. Gartner (17/58. Gartner 'Bastarderzeugung' s. 259, 518, 526 et seq.) sums up his conviction on this head in the strongest terms. Kolreuter (17/59. 'Fortsetzung' 1763 s. 29; 'Dritte Fortsetzung' s. 44, 96; 'Act. Acad. St. Petersburg' 1782 part 2 page 251; 'Nova Acta' 1793 pages 391, 394; 'Nova Acta' 1795 pages 316, 323.) gives numerous precise measurements of the weight and height of his hybrids in his comparison with measurements of both parent-forms; and speaks with astonishment of their "statura portentosa," their "ambitus vastissimus ac altitudo valde conspicua." Some exceptions to the rule in the case of very sterile hybrids have, however, been noticed by Gartner and Herbert; but the most striking exceptions are given by Max Wichura (17/60.

Charles Darwin

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