Hence it would appear probable, that the same causes which here make the immigrants of some peculiar species smaller, make most of the peculiar Galapageian species also smaller, as well as very generally more dusky coloured. All the plants have a wretched, weedy appearance, and I did not see one beautiful flower. The insects, again, are small-sized and dull-coloured, and, as Mr. Waterhouse informs me, there is nothing in their general appearance which would have led him to imagine that they had come from under the equator. [1] The birds, plants, and insects have a desert character, and are not more brilliantly coloured than those from southern Patagonia; we may, therefore, conclude that the usual gaudy colouring of the inter-tropical productions, is not related either to the heat or light of those zones, but to some other cause, perhaps to the conditions of existence being generally favourable to life.

We will now turn to the order of reptiles, which gives the most striking character to the zoology of these islands. The species are not numerous, but the numbers of individuals of each species are extraordinarily great. There is one small lizard belonging to a South American genus, and two species (and probably more) of the Amblyrhynchus -- a genus confined to the Galapagos Islands. There is one snake which is numerous; it is identical, as I am informed by M. Bibron, with the Psammophis Temminckii from Chile. [2] Of sea- turtle I believe there are more than one species, and of tortoises there are, as we shall presently show, two or three species or races. Of toads and frogs there are none: I was surprised at this, considering how well suited for them the temperate and damp upper woods appeared to be. It recalled to my mind the remark made by Bory St. Vincent, [3] namely, that none of this family are found on any of the volcanic islands in the great oceans. As far as I can ascertain from various works, this seems to hold good throughout the Pacific, and even in the large islands of the Sandwich archipelago. Mauritius offers an apparent exception, where I saw the Rana Mascariensis in abundance: this frog is said now to inhabit the Seychelles, Madagascar, and Bourbon; but on the other hand, Du Bois, in his voyage in 1669, states that there were no reptiles in Bourbon except tortoises; and the Officier du Roi asserts that before 1768 it had been attempted, without success, to introduce frogs into Mauritius -- I presume for the purpose of eating: hence it may be well doubted whether this frog is an aboriginal of these islands. The absence of the frog family in the oceanic islands is the more remarkable, when contrasted with the case of lizards, which swarm on most of the smallest islands. May this difference not be caused, by the greater facility with which the eggs of lizards, protected by calcareous shells might be transported through salt-water, than could the slimy spawn of frogs?

I will first describe the habits of the tortoise (Testudo nigra, formerly called Indica), which has been so frequently alluded to. These animals are found, I believe, on all the islands of the archipelago; certainly on the greater number. They frequent in preference the high damp parts, but they likewise live in the lower and arid districts. I have already shown, from the numbers which have been caught in a single day, how very numerous they must be. Some grow to an immense size: Mr. Lawson, an Englishman, and vice-governor of the colony, told us that he had seen several so large, that it required six or eight men to lift them from the ground; and that some had afforded as much as two hundred pounds of meat. The old males are the largest, the females rarely growing to so great a size: the male can readily be distinguished from the female by the greater length of its tail. The tortoises which live on those islands where there is no water, or in the lower and arid parts of the others, feed chiefly on the succulent cactus. Those which frequent the higher and damp regions, eat the leaves of various trees, a kind of berry (called guayavita) which is acid and austere, and likewise a pale green filamentous lichen (Usnera plicata), that hangs from the boughs of the trees.

The tortoise is very fond of water, drinking large quantities, and wallowing in the mud. The larger islands alone possess springs, and these are always situated towards the central parts, and at a considerable height. The tortoises, therefore, which frequent the lower districts, when thirsty, are obliged to travel from a long distance. Hence broad and well-beaten paths branch off in every direction from the wells down to the sea-coast; and the Spaniards by following them up, first discovered the watering-places. When I landed at Chatham Island, I could not imagine what animal travelled so methodically along well-chosen tracks. Near the springs it was a curious spectacle to behold many of these huge creatures, one set eagerly travelling onwards with outstretched necks, and another set returning, after having drunk their fill. When the tortoise arrives at the spring, quite regardless of any spectator, he buries his head in the water above his eyes, and greedily swallows great mouthfuls, at the rate of about ten in a minute. The inhabitants say each animal stays three or four days in the neighbourhood of the water, and then returns to the lower country; but they differed respecting the frequency of these visits. The animal probably regulates them according to the nature of the food on which it has lived. It is, however, certain, that tortoises can subsist even on these islands where there is no other water than what falls during a few rainy days in the year.

Charles Darwin

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