Will you, therefore, look over the few pages which I have sent, and tell me whether you find any flaw, or whether you think I should change the form of expression? You have been so unhandsomely and uncandidly dealt with by a friend of yours and mine that I should be sorry to find myself in the position of an opponent to you, and more particularly with the chance of making a fool of myself.
I met your brother yesterday, who tells me you are coming to town. I hope you will give me a hail. I long for a jaw with you, and have much to speak to you about.
You will have seen the eclaircissement about the Eocene monkeys of England. By a touch of the conjuring wand they have been metamorphosed--a la Darwin --into Hyracotherian pigs. (142/2. "On the Hyracotherian Character of the Lower Molars of the supposed Macacus from the Eocene Sand of Kyson, Suffolk." "Ann. Mag. Nat. Hist." Volume X., 1862, page 240. In this note Owen stated that the teeth which he had named Macacus ("Ann. Mag." 1840, page 191) most probably belonged to Hyracotherium cuniculus. See "A Catalogue of British Fossil Vertebrata," A.S. Woodward and C.D. Sherborn, 1890, under Hyracotherium, page 356; also Zittel's "Handbuch der Palaeontologie" Abth. I., Bd. IV., Leipzig, 1891-93, page 703.) Would you believe it? This even is a gross blunder. They are not pigs.
LETTER 143. TO HUGH FALCONER. Down, October 1st [1862].
On my return home yesterday I found your letter and MS., which I have read with extreme interest. Your note and every word in your paper are expressed with the same kind feeling which I have experienced from you ever since I have had the happiness of knowing you. I value scientific praise, but I value incomparably higher such kind feeling as yours. There is not a single word in your paper to which I could possibly object: I should be mad to do so; its only fault is perhaps its too great kindness. Your case seems the most striking one which I have met with of the persistence of specific characters. It is very much the more striking as it relates to the molar teeth, which differ so much in the species of the genus, and in which consequently I should have expected variation. As I read on I felt not a little dumbfounded, and thought to myself that whenever I came to this subject I should have to be savage against myself; and I wondered how savage you would be. I trembled a little. My only hope was that something could be made out of the bog N. American forms, which you rank as a geographical race; and possibly hereafter out of the Sicilian species. Guess, then, my satisfaction when I found that you yourself made a loophole (143/1. This perhaps refers to a passage ("N.H. Review," 1863, page 79) in which Falconer allows the existence of intermediate forms along certain possible lines of descent. Falconer's reference to the Sicilian elephants is in a note on page 78; the bog-elephant is mentioned on page 79.), which I never, of course, could have guessed at; and imagine my still greater satisfaction at your expressing yourself as an unbeliever in the eternal immutability of species. Your final remarks on my work are too generous, but have given me not a little pleasure. As for criticisms, I have only small ones. When you speak of "moderate range of variation" I cannot but think that you ought to remind your readers (though I daresay previously done) what the amount is, including the case of the American bog-mammoth. You speak of these animals as having been exposed to a vast range of climatal changes from before to after the Glacial period. I should have thought, from analogy of sea-shells, that by migration (or local extinction when migration not possible) these animals might and would have kept under nearly the same climate.
A rather more important consideration, as it seems to me, is that the whole proboscidean group may, I presume, be looked at as verging towards extinction: anyhow, the extinction has been complete as far as Europe and America are concerned. Numerous considerations and facts have led me in the "Origin" to conclude that it is the flourishing or dominant members of each order which generally give rise to new races, sub-species, and species; and under this point of view I am not at all surprised at the constancy of your species.