If the referee on my paper should recommend it to appear in the "Transactions," will you be so kind as to lay my earnest request before the Council that it may be permitted to appear in the "Journal?"
You must be very busy with your change of residence; but when you are settled and have some leisure, perhaps you will be so kind as to give me some cases of dimorphism, like that of Primula. Should you object to my adding them to those given me by A. Gray? By the way, I heard from A. Gray this morning, and he gives me two very curious cases in Boragineae.
LETTER 608. TO JOHN LINDLEY.
(608/1. In the following fragment occurs the earliest mention of Darwin's work on the three sexual forms of Catasetum tridentatum. Sir R. Schomburgk (608/2. "Trans. Linn. Soc." XVII., page 522.) described Catasetum tridentatum, Monacanthus viridis and Myanthus barbatus occurring on a single plant, but it remained for Darwin to make out that they are the male, female and hermaphrodite forms of a single species. (608/3. "Fertilisation of Orchids," Edition I., page 236; Edition II., page 196.)
With regard to the species of Acropera (Gongora) (608/4. Acropera Loddigesii = Gongora galeata: A. luteola = G. fusca ("Index Kewensis").) he was wrong in his surmise. The apparent sterility seems to be explicable by Hildebrand's discovery (608/5. "Bot. Zeitung," 1863 and 1865.) that in some orchids the ovules are not developed until pollinisation has occurred. (608/6. "Fertilisation of Orchids," Edition II., page 172. See Letter 633.))
Down, December 15th [1861].
I am so nearly ready for press that I will not ask for anything more; unless, indeed, you stumbled on Mormodes in flower. As I am writing I will just mention that I am convinced from the rudimentary state of the ovules, and from the state of the stigma, that the whole plant of Acropera luteola (and I believe A. Loddigesii) is male. Have you ever seen any form from the same countries which could be the females? Of course no answer is expected unless you have ever observed anything to bear on this. I may add [judging from the] state of the ovules and of the pollen [that]:--
Catasetum tridentatum is male (and never seeds, according to Schomburgk, whom you have accidentally misquoted in the "Vegetable Kingdom"). Monacanthus viridis is female. Myanthus barbatus is the hermaphrodite form of same species.
LETTER 609. TO J.D. HOOKER. Down, December 18th [1861].
Thanks for your note. I have not written for a long time, for I always fancy, busy as you are, that my letters must be a bore; though I like writing, and always enjoy your notes. I can sympathise with you about fear of scarlet fever: to the day of my death I shall never forget all the sickening fear about the other children, after our poor little baby died of it. The "Genera Plantarum" must be a tremendous work, and no doubt very valuable (such a book, odd as it may appear, would be very useful even to me), but I cannot help being rather sorry at the length of time it must take, because I cannot enter on and understand your work. Will you not be puzzled when you come to the orchids? It seems to me orchids alone would be work for a man's lifetime; I cannot somehow feel satisfied with Lindley's classification; the Malaxeae and Epidendreae seem to me very artificially separated. (609/1. Pfitzer (in the "Pflanzenfamilien") places Epidendrum in the Laeliinae-Cattleyeae, Malaxis in the Liparidinae. He states that Bentham united the Malaxideae and Epidendreae.) Not that I have seen enough to form an opinion worth anything.
Your African plant seems to be a vegetable Ornithorhynchus, and indeed much more than that. (609/2. See Sir J.D. Hooker, "On Welwitschia, a new genus of Gnetaceae." "Linn. Soc. Trans." XXIV., 1862-3.) The more I read about plants the more I get to feel that all phanerogams seem comparable with one class, as lepidoptera, rather than with one kingdom, as the whole insecta. (609/3. He wrote to Hooker (December 28th, 1861): "I wrote carelessly about the value of phanerogams; what I was thinking of was that the sub- groups seemed to blend so much more one into another than with most classes of animals.