I have had a splendid day's work, and must tell you about it.
Lady Dorothy sent me a young plant of U[tricularia] montana (727/1. See "Life and Letters," III., page 327, and "Insectivorous Plants," page 431.), which I fancy is the species you told me of. The roots or rhizomes (for I know not which they are; I can see no scales or internodes or absorbent hairs) bear scores of bladders from 1/20 to 1/100 of an inch in diameter; and I traced these roots to the depth of 1 1/2 in. in the peat and sand. The bladders are like glass, and have the same essential structure as those of our species, with the exception that many exterior parts are aborted. Internally the structure is perfect, as is the minute valvular opening into the bladder, which is filled with water. I then felt sure that they captured subterranean insects, and after a time I found two with decayed remnants, with clear proof that something had been absorbed, which had generated protoplasm. When you are here I shall be very curious to know whether they are roots or rhizomes.
Besides the bladders there are great tuber-like swellings on the rhizomes; one was an inch in length and half in breadth. I suppose these must have been described. I strongly suspect that they serve as reservoirs for water. (727/2. The existence of water-stores is quite in accordance with the epiphytic habit of the plant.) But I shall experimentise on this head. A thin slice is a beautiful object, and looks like coarsely reticulated glass.
If you have an old plant which could be turned out of its pot (and can spare the time), it would be a great gain to me if you would tear off a bit of the roots near the bottom, and shake them well in water, and see whether they bear these minute glass-like bladders. I should also much like to know whether old plants bear the solid bladder-like bodies near the upper surface of the pot. These bodies are evidently enlargements of the roots or rhizomes. You must forgive this long letter, and make allowance for my delight at finding this new sub-group of insect-catchers. Sir E. Tennent speaks of an aquatic species of Utricularia in Ceylon, which has bladders on its roots, and rises annually to the surface, as he says, by this means. (727/3. Utricularia stellaris. Emerson Tennent's "Ceylon," Volume I., page 124, 1859.)
We shall be delighted to see you here on the 26th; if you will let us know your train we will send to meet you. You will have to work like a slave while you are here.
LETTER 728. TO J. JENNER WEIR.
(728/1. In 1870 Mr. Jenner Weir wrote to Darwin: "My brother has but two kinds of laburnum, viz., Cytisus purpureus, very erect, and Cytisus alpinus, very pendulous. He has several stocks of the latter grafted with the purple one; and this year, the grafts being two years old, I saw in one, fairly above the stock, about four inches, a raceme of purely yellow flowers with the usual dark markings, and above them a bunch of purely purple flowers; the branches of the graft in no way showed an intermediate character, but had the usual rigid growth of purpureus."
Early in July 1875, when Darwin was correcting a new edition of "Variation under Domestication," he again corresponded with Mr. Weir on the subject.)
Down, July 8th [1875].
I thank you cordially. The case interests me in a higher degree than anything which I have heard for a very long time. Is it your brother Harrison W., whom I know? I should like to hear where the garden is. There is one other very important point which I am most anxious to hear--viz., the nature of the leaves at the base of the yellow racemes, for leaves are always there produced with the yellow laburnums, and I suppose so in the case of C. purpureus. As the tree has produced yellow racemes several times, do you think you could ask your brother to cut off and send me by post in a box a small branch of the purple stock with the pods or leaves of the yellow sport? (728/2. "The purple stock" here means the supposed C. purpureus, on which a yellow-flowered branch was borne.) This would be an immense favour, for then I would cut the point of junction longitudinally and examine slice under the microscope, to be able to state no trace of bud of yellow kind having been inserted. I do not suspect anything of the kind, but it is sure to be said that your brother's gardener, either by accident or fraud, inserted a bud. Under this point of view it would be very good to gather from your brother how many times the yellow sport has appeared.