(55. Livingstone's 'Travels and Researches in S. Africa,' 1857, pp. 338, 339. D'Orbigny, as quoted by Godron, 'De l'Espece,' vol. ii. p. 266.)

Various facts, which I have given elsewhere, prove that the colour of the skin and hair is sometimes correlated in a surprising manner with a complete immunity from the action of certain vegetable poisons, and from the attacks of certain parasites. Hence it occurred to me, that negroes and other dark races might have acquired their dark tints by the darker individuals escaping from the deadly influence of the miasma of their native countries, during a long series of generations.

I afterwards found that this same idea had long ago occurred to Dr. Wells. (56. See a paper read before the Royal Soc. in 1813, and published in his Essays in 1818. I have given an account of Dr. Wells' views in the Historical Sketch (p. xvi.) to my 'Origin of Species.' Various cases of colour correlated with constitutional peculiarities are given in my 'Variation of Animals and Plants under Domestication,' vol. ii. pp. 227, 335.) It has long been known that negroes, and even mulattoes, are almost completely exempt from the yellow-fever, so destructive in tropical America. (57. See, for instance, Nott and Gliddon, 'Types of Mankind,' p. 68.) They likewise escape to a large extent the fatal intermittent fevers, that prevail along at least 2600 miles of the shores of Africa, and which annually cause one-fifth of the white settlers to die, and another fifth to return home invalided. (58. Major Tulloch, in a paper read before the Statistical Society, April 20, 1840, and given in the 'Athenaeum,' 1840, p. 353.) This immunity in the negro seems to be partly inherent, depending on some unknown peculiarity of constitution, and partly the result of acclimatisation. Pouchet (59. 'The Plurality of the Human Race' (translat.), 1864, p. 60.) states that the negro regiments recruited near the Soudan, and borrowed from the Viceroy of Egypt for the Mexican war, escaped the yellow-fever almost equally with the negroes originally brought from various parts of Africa and accustomed to the climate of the West Indies. That acclimatisation plays a part, is shewn by the many cases in which negroes have become somewhat liable to tropical fevers, after having resided for some time in a colder climate. (60. Quatrefages, 'Unite de l'Espece Humaine,' 1861, p. 205. Waitz, 'Introduction to Anthropology,' translat., vol. i. 1863, p. 124. Livingstone gives analogous cases in his 'Travels.') The nature of the climate under which the white races have long resided, likewise has some influence on them; for during the fearful epidemic of yellow fever in Demerara during 1837, Dr. Blair found that the death-rate of the immigrants was proportional to the latitude of the country whence they had come. With the negro the immunity, as far as it is the result of acclimatisation, implies exposure during a prodigious length of time; for the aborigines of tropical America who have resided there from time immemorial, are not exempt from yellow fever; and the Rev. H.B. Tristram states, that there are districts in Northern Africa which the native inhabitants are compelled annually to leave, though the negroes can remain with safety.

That the immunity of the negro is in any degree correlated with the colour of his skin is a mere conjecture: it may be correlated with some difference in his blood, nervous system, or other tissues. Nevertheless, from the facts above alluded to, and from some connection apparently existing between complexion and a tendency to consumption, the conjecture seemed to me not improbable. Consequently I endeavoured, with but little success (61. In the spring of 1862 I obtained permission from the Director-General of the Medical department of the Army, to transmit to the surgeons of the various regiments on foreign service a blank table, with the following appended remarks, but I have received no returns. "As several well-marked cases have been recorded with our domestic animals of a relation between the colour of the dermal appendages and the constitution; and it being notorious that there is some limited degree of relation between the colour of the races of man and the climate inhabited by them; the following investigation seems worth consideration.

Charles Darwin

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