86.), and, as we have seen, with the manes of some species of baboons. But with most kinds of monkeys the various tufts of hair about the face and head are alike in both sexes.
The males of various members of the ox family (Bovidae), and of certain antelopes, are furnished with a dewlap, or great fold of skin on the neck, which is much less developed in the female.
Now, what must we conclude with respect to such sexual differences as these? No one will pretend that the beards of certain male goats, or the dewlaps of the bull, or the crests of hair along the backs of certain male antelopes, are of any use to them in their ordinary habits. It is possible that the immense beard of the male Pithecia, and the large beard of the male orang, may protect their throats when fighting; for the keepers in the Zoological Gardens inform me that many monkeys attack each other by the throat; but it is not probable that the beard has been developed for a distinct purpose from that served by the whiskers, moustache, and other tufts of hair on the face; and no one will suppose that these are useful as a protection. Must we attribute all these appendages of hair or skin to mere purposeless variability in the male? It cannot be denied that this is possible; for in many domesticated quadrupeds, certain characters, apparently not derived through reversion from any wild parent form, are confined to the males, or are more developed in them than in the females-- for instance, the hump on the male zebu-cattle of India, the tail of fat- tailed rams, the arched outline of the forehead in the males of several breeds of sheep, and lastly, the mane, the long hairs on the hind legs, and the dewlap of the male of the Berbura goat. (18. See the chapters on these several animals in vol. i. of my 'Variation of Animals under Domestication;' also vol. ii. p. 73; also chap. xx. on the practice of selection by semi-civilised people. For the Berbura goat, see Dr. Gray, 'Catalogue,' ibid. p. 157.) The mane, which occurs only in the rams of an African breed of sheep, is a true secondary sexual character, for, as I hear from Mr. Winwood Reade, it is not developed if the animal be castrated. Although we ought to be extremely cautious, as shewn in my work on 'Variation under Domestication,' in concluding that any character, even with animals kept by semi-civilised people, has not been subjected to selection by man, and thus augmented, yet in the cases just specified this is improbable; more especially as the characters are confined to the males, or are more strongly developed in them than in the females. If it were positively known that the above African ram is a descendant of the same primitive stock as the other breeds of sheep, and if the Berbura male-goat with his mane, dewlap, etc., is descended from the same stock as other goats, then, assuming that selection has not been applied to these characters, they must be due to simple variability, together with sexually- limited inheritance.
Hence it appears reasonable to extend this same view to all analogous cases with animals in a state of nature. Nevertheless I cannot persuade myself that it generally holds good, as in the case of the extraordinary development of hair on the throat and fore-legs of the male Ammotragus, or in that of the immense beard of the male Pithecia. Such study as I have been able to give to nature makes me believe that parts or organs which are highly developed, were acquired at some period for a special purpose. With those antelopes in which the adult male is more strongly-coloured than the female, and with those monkeys in which the hair on the face is elegantly arranged and coloured in a diversified manner, it seems probable that the crests and tufts of hair were gained as ornaments; and this I know is the opinion of some naturalists. If this be correct, there can be little doubt that they were gained or at least modified through sexual selection; but how far the same view may be extended to other mammals is doubtful.