I have continued steadily reading and collecting facts on variation of domestic animals and plants, and on the question of what are species. I have a grand body of facts, and I think I can draw some sound conclusions. The general conclusions at which I have slowly been driven from a directly opposite conviction, is that species are mutable, and that allied species are co-descendants from common stocks. I know how much I open myself to reproach for such a conclusion, but I have at least honestly and deliberately come to it. I shall not publish on this subject for several years. At present I am on the Geology of South America. I hope to pick up from your book some facts on slight variations in structure or instincts in the animals of your acquaintance.
Believe me, ever yours, C. DARWIN.
CHARLES DARWIN TO L. JENYNS (REV. L. BLOMEFIELD). Down, [1845?].
My dear Jenyns,
I am very much obliged to you for the trouble you have taken in having written me so long a note. The question of where, when, and how the check to the increase of a given species falls appears to me particularly interesting, and our difficulty in answering it shows how really ignorant we are of the lives and habits of our most familiar species. I was aware of the bare fact of old birds driving away their young, but had never thought of the effect you so clearly point out, of local gaps in number being thus immediately filled up. But the original difficulty remains; for if your farmers had not killed your sparrows and rooks, what would have become of those which now immigrate into your parish? in the middle of England one is too far distant from the natural limits of the rook and sparrow to suppose that the young are thus far expelled from Cambridgeshire. The check must fall heavily at some time of each species' life; for, if one calculates that only half the progeny are reared and bred, how enormous is the increase! One has, however, no business to feel so much surprise at one's ignorance, when one knows how impossible it is without statistics to conjecture the duration of life and percentage of deaths to births in mankind. If it could be shown that apparently the birds of passage WHICH BREED HERE and increase, return in the succeeding years in about the same number, whereas those that come here for their winter and non-breeding season annually, come here with the same numbers, but return with greatly decreased numbers, one would know (as indeed seems probable) that the check fell chiefly on full-grown birds in the winter season, and not on the eggs and very young birds, which has appeared to me often the most probable period. If at any time any remarks on this subject should occur to you, I should be most grateful for the benefit of them.
With respect to my far distant work on species, I must have expressed myself with singular inaccuracy if I led you to suppose that I meant to say that my conclusions were inevitable. They have become so, after years of weighing puzzles, to myself ALONE; but in my wildest day-dream, I never expect more than to be able to show that there are two sides to the question of the immutability of species, i.e. whether species are DIRECTLY created or by intermediate laws (as with the life and death of individuals). I did not approach the subject on the side of the difficulty in determining what are species and what are varieties, but (though, why I should give you such a history of my doings it would be hard to say) from such facts as the relationship between the living and extinct mammifers in South America, and between those living on the Continent and on adjoining islands, such as the Galapagos. It occured to me that a collection of all such analogous facts would throw light either for or against the view of related species being co-descendants from a common stock. A long searching amongst agricultural and horticultural books and people makes me believe (I well know how absurdly presumptuous this must appear) that I see the way in which new varieties become exquisitely adapted to the external conditions of life and to other surrounding beings.