Within nearly this same period (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow- horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are. The more I reflect on this case, the more interesting it appears: I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well- characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earth's crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals [6] seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.
When America, and especially North America, possessed its elephants, mastodons, horse, and hollow-horned ruminants, it was much more closely related in its zoological characters to the temperate parts of Europe and Asia than it now is. As the remains of these genera are found on both sides of Behring's Straits [7] and on the plains of Siberia, we are led to look to the north-western side of North America as the former point of communication between the Old and so-called New World. And as so many species, both living and extinct, of these same genera inhabit and have inhabited the Old World, it seems most probable that the North American elephants, mastodons, horse, and hollow- horned ruminants migrated, on land since submerged near Behring's Straits, from Siberia into North America, and thence, on land since submerged in the West Indies, into South America, where for a time they mingled with the forms characteristic of that southern continent, and have since become extinct.
While travelling through the country, I received several vivid descriptions of the effects of a late great drought; and the account of this may throw some light on the cases where vast numbers of animals of all kinds have been embedded together. The period included between the years 1827 and 1830 is called the "gran seco," or the great drought. During this time so little rain fell, that the vegetation, even to the thistles, failed; the brooks were dried up, and the whole country assumed the appearance of a dusty high road. This was especially the case in the northern part of the province of Buenos Ayres and the southern part of St. Fe. Very great numbers of birds, wild animals, cattle, and horses perished from the want of food and water. A man told me that the deer [8] used to come into his courtyard to the well, which he had been obliged to dig to supply his own family with water; and that the partridges had hardly strength to fly away when pursued. The lowest estimation of the loss of cattle in the province of Buenos Ayres alone, was taken at one million head. A proprietor at San Pedro had previously to these years 20,000 cattle; at the end not one remained. San Pedro is situated in the middle of the finest country; and even now abounds again with animals; yet during the latter part of the "gran seco," live cattle were brought in vessels for the consumption of the inhabitants. The animals roamed from their estancias, and, wandering far southward, were mingled together in such multitudes, that a government commission was sent from Buenos Ayres to settle the disputes of the owners. Sir Woodbine Parish informed me of another and very curious source of dispute; the ground being so long dry, such quantities of dust were blown about, that in this open country the landmarks became obliterated, and people could not tell the limits of their estates.
I was informed by an eye-witness that the cattle in herds of thousands rushed into the Parana, and being exhausted by hunger they were unable to crawl up the muddy banks, and thus were drowned. The arm of the river which runs by San Pedro was so full of putrid carcasses, that the master of a vessel told me that the smell rendered it quite impassable. Without doubt several hundred thousand animals thus perished in the river: their bodies when putrid were seen floating down the stream; and many in all probability were deposited in the estuary of the Plata. All the small rivers became highly saline, and this caused the death of vast numbers in particular spots; for when an animal drinks of such water it does not recover. Azara describes [9] the fury of the wild horses on a similar occasion, rushing into the marshes, those which arrived first being overwhelmed and crushed by those which followed. He adds that more than once he has seen the carcasses of upwards of a thousand wild horses thus destroyed. I noticed that the smaller streams in the Pampas were paved with a breccia of bones but this probably is the effect of a gradual increase, rather than of the destruction at any one period. Subsequently to the drought of 1827 to 1832, a very rainy season followed which caused great floods. Hence it is almost certain that some thousands of the skeletons were buried by the deposits of the very next year. What would be the opinion of a geologist, viewing such an enormous collection of bones, of all kinds of animals and of all ages, thus embedded in one thick earthy mass? Would he not attribute it to a flood having swept over the surface of the land, rather than to the common order of things? [10]