Eyton.

SKULL.

(FIGURE 33. OCCIPITAL FORAMEN, of natural size. A. Wild Gallus bankiva. B. Cochin Cock.

FIGURE 34. SKULLS of natural size, viewed from above, a little obliquely. A. Wild Gallus bankiva. B. White-crested Polish Cock.

FIGURE 35. LONGITUDINAL SECTIONS OF SKULL, of natural size, viewed laterally; A. Polish Cock. B. Cochin Cock, selected for comparison with the above from being of nearly the same size.

FIGURE 36. SKULL OF HORNED FOWL, of natural size, viewed from above, a little obliquely. (In the possession of Tegetmeier.))

[The SKULL differs greatly in size in different breeds, being nearly twice as long in the largest Cochins, but not nearly twice as broad, as in Bantams. The bones at the base, from the occipital foramen to the anterior end (including the quadrates and pterygoids), are absolutely identical in SHAPE in all the skulls. So is the lower jaw. In the forehead slight differences are often perceptible between the males and females, evidently caused by the presence of the comb. In every case I take the skull of G. bankiva as the standard of comparison. In four Games, in one Malay hen, in an African cock, in a Frizzled cock from Madras, in two black-boned Silk hens, no differences worth notice occur. In three SPANISH cocks, the form of the forehead between the orbits differs considerably; in one it is considerably depressed, whilst in the two others it is rather prominent, with a deep medial furrow; the skull of the hen is smooth. In three skulls of SEBRIGHT BANTAMS the crown is more globular, and slopes more abruptly to the occiput, than in G. bankiva. In a Bantam or Jumper from Burmah these same characters are more strongly pronounced, and the supra-occiput is more pointed. In a black Bantam the skull is not so globular, and the occipital foramen is very large, and has nearly the same sub-triangular outline presently to be described in Cochins; and in this skull the two ascending branches of the premaxillary are overlapped in a singular manner by the processes of the nasal bone, but, as I have seen only one specimen, some of these differences may be individual. Of Cochins and Brahmas (the latter a crossed race approaching closely to Cochins) I have examined seven skulls; at the point where the ascending branches of the premaxillary rest on the frontal bone the surface is much depressed, and from this depression a deep medial furrow extends backwards to a variable distance; the edges of this fissure are rather prominent, as is the top of the skull behind and over the orbits. These characters are less developed in the hens. The pterygoids, and the processes of the lower jaw, are broader, relatively to the size of the head, than in G. bankiva; and this is likewise the case with Dorkings when of large size. The fork of the hyoid bone in Cochins is twice as wide as in G. bankiva, whereas the length of the other hyoid bones is only as three to two. But the most remarkable character is the shape of the occipital foramen: in G. bankiva (A) the breadth in a horizontal line exceeds the height in a vertical line, and the outline is nearly circular; whereas in Cochins (B) the outline is sub-triangular, and the vertical line exceeds the horizontal line in length. This same form likewise occurs in the black Bantam above referred to, and an approach to it may be seen in some Dorkings, and in a slight degree in certain other breeds.

Of Dorkings I have examined three skulls, one belonging to the white-sub- breed; the one character deserving notice is the breadth of the frontal bones, which are moderately furrowed in the middle; thus in a skull which was less than once and a half the length of that of G. bankiva, the breadth between the orbits was exactly double. Of Hamburghs I have examined four skulls (male and female) of the pencilled sub-breed, and one (male) of the spangled sub-breed; the nasal bones stand remarkably wide apart, but in a variable degree; consequently narrow membrane-covered spaces are left between the tips of the two ascending branches of the pre-maxillary bones, which are rather short, and between these branches and the nasal bones.

Charles Darwin

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