Captain Hutton (8/65. 'Transact. Entomolog. Soc.' 3rd series volume 3 pages 143-173 and pages 295-331.) states that throughout the world at least six species have been domesticated; and he believes that the silk-moths reared in Europe belong to two or three species. This, however, is not the opinion of several capable judges who have particularly attended to the cultivation of this insect in France; and hardly accords with some facts presently to be given.

The common silk-moth (Bombyx mori) was brought to Constantinople in the sixth century, whence it was carried into Italy, and in 1494 into France. (8/66. Godron 'De l'Espece' 1859 tome 1 page 460. The antiquity of the silkworm in China is given on the authority of Stanislas Julien.) Everything has been favourable for the variation of this insect. It is believed to have been domesticated in China as long ago as 2700 B.C. It has been kept under unnatural and diversified conditions of life, and has been transported into many countries. There is reason to believe that the nature of the food given to the caterpillar influences to a certain extent the character of the breed. (8/67. See the remarks of Prof. Westwood, Gen. Hearsey and others at the meeting of the Entomolog. Soc. of London July 1861.) Disuse has apparently aided in checking the development of the wings. But the most important element in the production of the many now existing, much modified races, no doubt has been the close attention which has long been applied in many countries to every promising variation. The care taken in Europe in the selection of the best cocoons and moths for breeding is notorious (8/68. See for instance M. A. de Quatrefages 'Etudes sur les Maladies actuelles du Ver a Soie' 1859 page 101.), and the production of eggs is followed as a distinct trade in parts of France. I have made inquiries through Dr. Falconer, and am assured that in India the natives are equally careful in the process of selection. In China the production of eggs is confined to certain favourable districts, and the raisers are precluded by law from producing silk, so that their whole attention may be necessarily given up to this one object. (8/69. My authorities for the statements will be given in the chapter on Selection.)

[The following details on the differences between the several breeds are taken, when not stated to the contrary, from M. Robinet's excellent work (8/70. 'Manuel de l'Educateur de Vers a Soie' 1848.), which bears every sign of care and large experience. The EGGS in the different races vary in colour, in shape (being round, elliptic or oval), and in size. The eggs laid in June in the south of France, and in July in the central provinces, do not hatch until the following spring; and it is in vain, says M. Robinet, to expose them to a temperature gradually raised, in order that the caterpillar may be quickly developed. Yet occasionally, without any known cause, batches of eggs are produced, which immediately begin to undergo the proper changes, and are hatched in from twenty to thirty days. From these and some other analogous facts it may be concluded that the Trevoltini silkworms of Italy, of which the caterpillars are hatched in from fifteen to twenty days, do not necessarily form, as has been maintained, a distinct species. Although the breeds which live in temperate countries produce eggs which cannot be immediately hatched by artificial heat, yet when they are removed to and reared in a hot country they gradually acquire the character of quick development, as in the Trevoltini races. (8/71. Robinet ibid pages 12, 318. I may add that the eggs of N. American silkworms taken to the Sandwich Islands produced moths at very irregular periods; and the moths thus raised yielded eggs which were even worse in this respect. Some were hatched in ten days, and others not until after the lapse of many months. No doubt a regular early character would ultimately have been acquired. See review in 'Athenaeum' 1844 page 329 of J. Jarves 'Scenes in the Sandwich Islands.')

CATERPILLARS.

Charles Darwin

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