Maize has varied in an extraordinary and conspicuous manner. Metzger (9/55. 'Die Getreidearten' 1841 s. 208. I have modified a few of Metzger's statements in accordance with those made by Bonafous in his great work 'Hist. Nat. du Mais' 1836.), who paid particular attention to the cultivation of this plant, makes twelve races (unter-art) with numerous sub-varieties: of the latter some are tolerably constant, others quite inconstant. The different races vary in height from 15-18 feet to only 16- 18 inches, as in a dwarf variety described by Bonafous. The whole ear is variable in shape, being long and narrow, or short and thick, or branched. The ear in one variety is more than four times as long as in a dwarf kind. The seeds are arranged in the ear in from six to even twenty rows, or are placed irregularly. The seeds are coloured--white, pale-yellow, orange, red, violet, or elegantly streaked with black (9/56. Godron 'De l'Espece' tome 2 page 80; Al. De Candolle ibid page 951.); and in the same ear there are sometimes seeds of two colours. In a small collection I found that a single grain of one variety nearly equalled in weight seven grains of another variety. The shape of the seed varies greatly, being very flat, or nearly globular, or oval; broader than long, or longer than broad; without any point, or produced into a sharp tooth, and this tooth is sometimes recurved. One variety (the rugosa of Bonafous, and which is extensively cultivated in the United States as sweet corn) has its seeds curiously wrinkled, giving to the whole ear a singular appearance. Another variety (the cymosa of Bon.) carries its ears so crowded together that it is called mais a bouquet. The seeds of some varieties contain much glucose instead of starch. Male flowers sometimes appear amongst the female flowers, and Mr. J. Scott has lately observed the rarer case of female flowers on a true male panicle, and likewise hermaphrodite flowers. (9/57. 'Transact. Bot. Soc. of Edinburgh' volume 8 page 60.) Azara describes (9/58. 'Voyages dans l'Amerique Meridionale' tome 1 page 147.) a variety in Paraguay the grains of which are very tender, and he states that several varieties are fitted for being cooked in various ways. The varieties also differ greatly in precocity, and have different powers of resisting dryness and the action of violent wind. (9/59. Bonafous 'Hist. Nat. du Mais' page 31.) Some of the foregoing differences would certainly be considered of specific value with plants in a state of nature.

Le Comte Re states that the grains of all the varieties which he cultivated ultimately assumed a yellow colour. But Bonafous (9/60. Ibid page 31.) found that most of those which he sowed for ten consecutive years kept true to their proper tints; and he adds that in the valleys of the Pyrenees and on the plains of Piedmont a white maize has been cultivated for more than a century, and has undergone no change.

The tall kinds grown in southern latitudes, and therefore exposed to great heat, require from six to seven months to ripen their seed; whereas the dwarf kinds, grown in northern and colder climates, require only from three to four months. (9/61. Metzger 'Getreidearten' s. 206.) Peter Kalm (9/62. 'Description of Maize' by P. Kalm 1752 in 'Swedish Acts' volume 4. I have consulted an old English MS. translation.), who particularly attended to this plant, says, that in the United States, in proceeding from south to north, the plants steadily diminish in bulk. Seeds brought from lat. 37 deg in Virginia, and sown in lat. 43-44 deg in New England, produce plants which will not ripen their seed, or ripen them with the utmost difficulty. So it is with seed carried from New England to lat. 45-47 deg in Canada. By taking great care at first, the southern kinds after some years' culture ripen their seed perfectly in their northern homes, so that this is an analogous case with that of the conversion of summer into winter wheat, and conversely. When tall and dwarf maize are planted together, the dwarf kinds are in full flower before the others have produced a single flower; and in Pennsylvania they ripen their seeds six weeks earlier than the tall maize. Metzger also mentions a European maize which ripens its seed four weeks earlier than another European kind.

Charles Darwin

All Pages of This Book