(11/43. 'Gardener's Chronicle' 1845 page 564.) Although several moss-roses have thus certainly arisen by bud-variation, the greater number probably owe their origin to seed of moss-roses. For Mr. Rivers informs me that his seedlings from the old single moss-rose almost always produced moss-roses; and the old single moss-rose was, as we have seen, the product by bud-variation of the double moss-rose originally imported from Italy. That the original moss-rose was the product of bud-variation is probable, from the facts above given and from the de Meaux moss-rose (also a variety of R. centifolia) (11/44. 'Transact. Hort. Soc.' volume 2 page 242.) having appeared as a sporting branch on the common rose de Meaux. Prof. Caspary has carefully described (11/45. 'Shriften der Phys. Oekon. Gesell. zu Konigsberg' February 3, 1865 s. 4. See also Dr. Caspary's paper in 'Transactions of the Hort. Congress of Amsterdam' 1865.) the case of a six-year-old white moss-rose, which sent up several suckers, one of which was thorny, and produced red flowers, destitute of moss, exactly like those of the Provence rose (R. centifolia): another shoot bore both kinds of flowers, and in addition longitudinally striped flowers. As this white moss-rose had been grafted on the Provence rose, Prof. Caspary attributes the above changes to the influence of the stock; but from the facts already given, and from others to be given, bud- variation, with reversion, is probably a sufficient explanation.
Many other instances could be added of roses varying by buds. The white Provence rose apparently originated in this way. (11/46. 'Gardener's Chronicle' 1852 page 759.) M. Carriere states (page 36) that he himself knows of five varieties thus produced by the Baronne Prevost. The double and highly-coloured Belladonna rose has produced by suckers both semi- double and almost single white roses (11/47. 'Transact. Hort. Soc.' volume 2 page 242.); whilst suckers from one of these semi-double white roses reverted to perfectly characterised Belladonnas. In St. Domingo, varieties of the China rose propagated by cuttings often revert after a year or two into the old China rose. (11/48. Sir R. Schomburgk 'Proc. Linn. Soc. Bot.' volume 2 page 132.) Many cases have been recorded of roses suddenly becoming striped or changing their character by segments: some plants of the Comtesse de Chabrillant, which is properly rose-coloured, were exhibited in 1862 (11/49. 'Gardener's Chronicle' 1862 page 619.), with crimson flakes on a rose ground. I have seen the Beauty of Billiard with a quarter and with half the flower almost white. 'The Austrian bramble (R. lutea) not rarely (11/50. Hopkirk 'Flora Anomala' 167.) produces branches with pure yellow flowers; and Prof. Henslow has seen exactly half the flower of a pure yellow, and I have seen narrow yellow streaks on a single petal, of which the rest was of the usual copper colour.
The following cases are highly remarkable. Mr. Rivers, as I am informed by him, possessed a new French rose with delicate smooth shoots, pale glaucous-green leaves, and semi-double pale flesh-coloured flowers striped with dark red; and on branches thus characterised there suddenly appeared in more than one instance, the famous old rose called the Baronne Prevost, with its stout thorny shoots, and immense, uniformly and richly coloured double flowers; so that in this case the shoots, leaves, and flowers, all at once changed their character by bud-variation. According to M. Verlot (11/51. 'Sur La Production et la Fixation des Varietes' 1865 page 4.), a variety called Rosa cannabifolia, which has peculiarly shaped leaflets, and differs from every member of the family in the leaves being opposite instead of alternate, suddenly appeared on a plant of R. alba in the gardens of the Luxembourg. Lastly, "a running shoot" was observed by Mr. H. Curtis (11/52. 'Journal of Horticulture' March 1865 page 233.) on the old Aimee Vibert Noisette, and he budded it on Celine; thus a climbing Aimee Vibert was first produced and afterwards propagated.