I have seen the potatoes, and they are very curious."

We will now turn to the experiments made in Germany, since the publication of Prof. Hildebrand's paper. Herr Magnus relates (11/114. 'Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin' October 17, 1871.) the results of numerous trials made by Herren Reuter and Lindemuth, both attached to the Royal Gardens of Berlin. They inserted the eyes of red potatoes into white ones, and vice versa. Many different forms partaking of the characters of the inserted bud and of the stock were thus obtained; for instance, some of the tubers were white with red eyes.

Herr Magnus also exhibited in the following year before the same Society (November 19, 1872), the produce of grafts between black, white, and red potatoes, made by Dr. Neubert. These were made by uniting not the tubers but the young stems, as was done by Mr. Fitzpatrick. The result was remarkable, inasmuch as all the tubers thus produced were intermediate in character, though in a variable degree. Those between the black and the white or the red were the most striking in appearance. Some from between the white and red had one half of one colour and the other half of the other colour.

At the next meeting of the society Herr Magnus communicated the results of Dr. Heimann's experiments in grafting together the tubers of red Saxon, blue, and elongated white potatoes. The eyes were removed by a cylindrical instrument, and inserted into corresponding holes in other varieties. The plants thus produced yielded a great number of tubers, which were intermediate between the two parent-forms in shape, and in the colour both of the flesh and skin.

Herr Reuter experimented (11/115. Ibid November 17, 1874. See also excellent remarks by Herr Magnus.) by inserting wedges of the elongated White Mexican potato into a Black Kidney potato. Both sorts are known to be very constant, and differ much not only in form and colour, but in the eyes of the Black Kidney being deeply sunk, whereas those of the White Mexican are superficial and of a different shape. The tubers produced by these hybrids were intermediate in colour and form; and some which resembled in form the graft, i.e. the Mexican, had eyes deeply sunk and of the same shape as in the stock or Black Kidney.]

Any one who will attentively consider the abstract now given, of the experiments made by many observers in several countries, will, I think, be convinced that by grafting two varieties of the potato together in various ways, hybridised plants can be produced. It should be observed that several of the experimentalists are scientific horticulturists, and some of them potato-growers on a large scale, who, though beforehand sceptical, have been fully convinced of the possibility, even of the ease, of making graft- hybrids. The only way of escaping from this conclusion is to attribute all the many recorded cases to simple bud-variation. Undoubtedly the potato, as we have seen in this chapter, does sometimes, though not often, vary by buds; but it should be especially noted that it is experienced potato- growers, whose business it is to look out for new varieties, who have expressed unbounded astonishment at the number of new forms produced by graft-hybridisation. It may be argued that it is merely the operation of grafting, and not the union of two kinds, which causes so extraordinary an amount of bud-variation; but this objection is at once answered by the fact that potatoes are habitually propagated by the tubers being cut into pieces, and the sole difference in the case of graft-hybrids is that either a half or a smaller segment or a cylinder is placed in close opposition with the tissue of another variety. Moreover, in two cases, the young stems were grafted together, and the plants thus united yielded the same results as when the tubers were united. It is an argument of the greatest weight that when varieties are produced by simple bud-variation, they frequently present quite new characters; whereas in all the numerous cases above given, as Herr Magnus likewise insists, the graft-hybrids are intermediate in character between the two forms employed.

Charles Darwin

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