Paget, both the milk and the second teeth assume a peculiar and characteristic form. Professor Rolleston, also, informs me that the incisor teeth are sometimes furnished with a vascular rim in correlation with intra-pulmonary deposition of tubercles. In other cases of phthisis and of cyanosis the nails and finger- ends become clubbed like acorns. I believe that no explanation has been offered of these and of many other cases of correlated disease.

What can be more curious and less intelligible than the fact previously given, on the authority of Mr. Tegetmeier, that young pigeons of all breeds, which when mature have white, yellow, silver-blue, or dun-coloured plumage, come out of the egg almost naked; whereas pigeons of other colours when first born are clothed with plenty of down? White Pea-fowls, as has been observed both in England and France (25/34. Rev. E.S. Dixon 'Ornamental Poultry' 1848 page 111; Isidore Geoffroy 'Hist. Anomalies' tome 1 page 211.), and as I have myself seen, are inferior in size to the common coloured kind; and this cannot be accounted for by the belief that albinism is always accompanied by constitutional weakness; for white or albino moles are generally larger than the common kind.

To turn to more important characters: the niata cattle of the Pampas are remarkable from their short foreheads, upturned muzzles, and curved lower jaws. In the skull the nasal and premaxillary bones are much shortened, the maxillaries are excluded from any junction with the nasals, and all the bones are slightly modified, even to the plane of the occiput. From the analogous case of the dog, hereafter to be given, it is probable that the shortening of the nasal and adjoining bones is the proximate cause of the other modifications in the skull, including the upward curvature of the lower jaw, though we cannot follow out the steps by which these changes have been effected.

Polish fowls have a large tuft of feathers on their heads; and their skulls are perforated by numerous holes, so that a pin can be driven into the brain without touching any bone. That this deficiency of bone is in some way connected with the tuft of feathers is clear from tufted ducks and geese likewise having perforated skulls. The case would probably be considered by some authors as one of balancement or compensation. In the chapter on Fowls, I have shown that with Polish fowls the tuft of feathers was probably at first small; by continued selection it became larger, and then rested on a fibrous mass; and finally, as it became still larger, the skull itself became more and more protuberant until it acquired its present extraordinary structure. Through correlation with the protuberance of the skull, the shape and even the relative connection of the premaxillary and nasal bones, the shape of the orifice of the nostrils, the breadth of the frontal bone, the shape of the post-lateral processes of the frontal and squamosal bones, and the direction of the bony cavity of the ear, have all been modified. The internal configuration of the skull and the whole shape of the brain have likewise been altered in a truly marvellous manner.

After this case of the Polish fowl it would be superfluous to do more than refer to the details previously given on the manner in which the changed form of the comb has affected the skull, in various breeds of the fowl, causing by correlation crests, protuberances, and depressions on its surface.

With our cattle and sheep the horns stand in close connection with the size of the skull, and with the shape of the frontal bones; thus Cline (25/35. 'On the Breeding of Domestic Animals' 1829 page 6.) found that the skull of a horned ram weighed five times as much as that of a hornless ram of the same age. When cattle become hornless, the frontal bones are "materially diminished in breadth towards the poll;" and the cavities between the bony plates "are not so deep, nor do they extend beyond the frontals." (25/36. Youatt on 'Cattle' 1834 page 283.)

It may be well here to pause and observe how the effects of correlated variability, of the increased use of parts, and of the accumulation of so- called spontaneous variations through natural selection, are in many cases inextricably commingled.

Charles Darwin

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