It may be said that the lateral ducts in the labellum running into the antero-lateral ovarian bundle is no argument that the labellum consists of three organs blended together.
In desperation (and from the curious way the base of upper petals are soldered at basal edges) I fancied the real form of upper sepal, upper petal and lower sepal might be as represented by red lines, and that there had been an incredible amount of splitting of sepals and petals and subsequent fusion.
This seems a monstrous notion, but I have just looked at Bauer's drawing of allied Bonatea, and there is a degree of lobing of petals and sepals which would account for anything.
Now could you spare me a dry flower out of your Herbarium of Bonatea speciosa (605/5. See "Fertilisation of Orchids," Edition I., page 304 (note), where the resemblances between the anomalous vessels of Bonatea and Habenaria are described. On November 14th, 1861, he wrote to Sir Joseph: "You are a true friend in need. I can hardly bear to let the Bonatea soak long enough."), that I might soak and look for ducts. If I cannot explain the case of Habenaria all my work is smashed. I was a fool ever to touch orchids.
LETTER 606. TO J.D. HOOKER. Down, November 17th [1861].
What two very interesting and useful letters you have sent me. You rather astound me with respect to value of grounds of generalisation in the morphology of plants. It reminds me that years ago I sent you a grass to name, and your answer was, "It is certainly Festuca (so-and-so), but it agrees as badly with the description as most plants do." I have often laughed over this answer of a great botanist...Lindley, from whom I asked for an orchid with a simple labellum, has most kindly sent me a lot of what he marks "rare" and "rarissima" of peloric orchids, etc., but as they are dried I know not whether they will be of use. He has been most kind, and has suggested my writing to Lady D. Nevill, who has responded in a wonderfully kind manner, and has sent a lot of treasures. But I must stop; otherwise, by Jove, I shall be transformed into a botanist. I wish I had been one; this morphology is surprisingly interesting. Looking to your note, I may add that certainly the fifteen alternating bundles of spiral vessels (mingled with odd beadlike vessels in some cases) are present in many orchids. The inner whorl of anther ducts are oftenest aborted. I must keep clear of Apostasia, though I have cast many a longing look at it in Bauer. (606/1. Apostasia has two fertile anthers like Cypripedium. It is placed by Engler and Prantl in the Apostasieae or Apostasiinae, among the Orchideae, by others in a distinct but closely allied group.)
I hope I may be well enough to read my own paper on Thursday, but I have been very seedy lately. (606/2. "On the two Forms, or Dimorphic Condition, in the Species of the Genus Primula," "Linn. Soc. Journ." 1862. He did read the paper, but it cost him the next day in bed. "Life and Letters," III., page 299.) I see there is a paper at the Royal on the same night, which will more concern you, on fossil plants of Bovey (606/3. Oswald Heer, "The Fossil Flora of Bovey Tracey," "Phil. Trans. R. Soc." 1862, page 1039.), so that I suppose I shall not have you; but you must read my paper when published, as I shall very much like to hear what you think. It seems to me a large field for experiment. I shall make use of my Orchid little volume in illustrating modification of species doctrine, but I keep very, very doubtful whether I am not doing a foolish action in publishing. How I wish you would keep to your old intention and write a book on plants. (606/4. Possibly a book similar to that described in Letter 696.)
LETTER 607. TO G. BENTHAM. Down, November 26th [1861].
Our notes have crossed on the road. I know it is an honour to have a paper in the "Transactions," and I am much obliged to you for proposing it, but I should greatly prefer to publish in the "Journal." Nor does this apply exclusively to myself, for in old days at the Geological Society I always protested against an abstract appearing when the paper itself might appear. I abominate also the waste of time (and it would take me a day) in making an abstract.